During the Battle of Thermopylae, King Leonidas leads the Spartan army against a Persian charge. The battle is won, but Leonidas is informed by a captured enemy soldier that the existence of the mountain path has been revealed to the Persian army, which is moving to surround the Spartans by morning. Nevertheless, Leonidas resolves to hold off the Persian advance.
In the present, Layla Hassan recovers the Spear of Leonidas and together with Victoria Bibeau, extracts the DNA of two individuals from it, the siblings Kassandra and Alexios. With help from the Assassins, Layla picks one of the siblings (the “Misthios”) and activates the Animus to find the location of the Staff of Hermes.
The Misthios started as a young Spartan child, raised by their parents Nikolaos and Myrrine, and inherited the Spear of Leonidas from Myrrine as one of Leonidas’ descendants. However, one day, both the Misthios and their sibling are thrown off a mountain due to an oracle’s prophecy, with the Misthios being dropped by Nikolaos himself. The Misthios survived the fall and fled to the island of Kephallonia, where they grew up performing odd jobs until the Peloponnesian War begins.